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There has been a church dedicated to St Margaret of Antioch on this site for some 900 years. The fourth church was destroyed in the Great Fire of 1666 and rebuilt by Wren. St Margaret Pattens was so-called to distinguish it from other City churches dedicated to St Margaret, the name possibly originating from a benefactor, or more likely connected with 'pattens', a type of undershoe made and sold near here. Rood Lane was at one time called St Margaret Pattens, its name changing when a rood or cross was erected in the churchyard while the church was being rebuilt. The small paved area to the south of the church features a single tree, but is important in the transition from the medieval street to the modern street of Eastcheap.

The church of St Martin Orgar formerly stood here, its name arising from a Deacon called Ordgarus who owned the church, which he presented prior to c.1181 to the Dean and Chapter of St Paul's. It had a churchyard by c.1250. The building was damaged but not entirely destroyed in the Great Fire and part of the tower and nave were subsequently repaired and converted as a meeting place and chapel for French Huguenot Protestants. By c.1820 this had become ruinous and all but the tower was demolished, which acted as the entrance to the burial ground for the parish of St Clement and now occupied the site of the old church. The former churchyard remains as a raised garden in private use.

The parish of St Martin Outwich dates from c.1196, with a church first mentioned in 1217, erected at the cost of the Oteswich family, hence its name. In 1385 it was vested in the Merchant Taylors' Company. The church escaped the Great Fire of 1666 but was destroyed by a fire in 1765. Although the church was rebuilt in 1796-98, it was later demolished in 1874, after the parish was united with that of St Helen Bishopsgate in 1872. The churchyard was separate from the church on a site in Camomile Street, presented by Robert Hide in 1538, now marked by a raised shrub bed.

St Mary Aldermanbury may derive its name from the Aldermans' Berry or Court Hall, which stood nearby, or from a family by the name of Aldermanbir associated with the church. There was a church by at least 1181 and a churchyard by c1250. The old church was largely destroyed in the Great Fire of 1666 and rebuilt by Christopher Wren. Following WWII bomb damage, the church and churchyard site was acquired by the Corporation of London with residual land from Wood Street Police Station, and laid out as public gardens. The footings of the C15th church, over which Wren had built, remain in the lawn in the east of the garden. To the west is a monument to John Heminge and Henry Condell, fellow actors of Shakespeare, after whose death in 1616 they collected his works and published them at their own personal expense.

St Mary Aldermary is an early church, described by John Stow as 'older than any church of St Mary in the Citie' and Aldermary (the elder Mary) may reflect this. Various additions to the building took place over the years, including partial rebuilding in 1510/11 at the behest of the Lord Mayor of London, Sir Henry Keble. In 1534 one of its rectors, Henry Gold, was executed for his involvement with the 'Holy Maid of Kent'. John Milton married his third wife here in 1662. Damaged in the Great Fire of 1666, the church was rebuilt by Christopher Wren. The former churchyard is now laid out as a small garden with a central tree, three headstones set against the wall among shrubbery and gravestones form a path leading to the west entrance of the church.

The church of St Mary at Hill is referred to in the late C12th and there was a churchyard on the north side by 1299. A number of former Lord Mayors of the C14th to C16th were buried here. The medieval church was partially destroyed in 1666 and rebuilt by Wren in 1670-74 re-using the medieval west end and heightening the north and south walls of the earlier building, much of which had survived. The churchyard was closed to burials in 1846 and it is simply laid out as a public garden, with paving stones, a central rectangular flower bed and a small tree; a lead rainwater butt (dated 1788) is used as a planter and two seats were provided by the MPGA. It is surrounded by a brick retaining wall.

In 1189 there is a reference to the church here as 'Ecclesia de Staningehage', Staining probably referring to a landowner from Staines. Oat Lane is probably so-called for an oat market that existed here by the C16th. St Mary Staining Church was destroyed in the Great Fire of 1666 and not restored. In 1965 the Corporation of London took over the maintenance of the open space created on the site of the church and churchyard. The garden is a raised area of lawn with a large plane tree, path on two sides and tombstones ranged along the back. It is overlooked by Pewterers' Hall, which was built here in 1959-61 after the Company's hall in Lime Street was demolished in 1932.

The site may once have been that of an ancient pre-Christian temple, fragments of which were discovered in 1716 when St Mary's was rebuilt. Possibly named after someone called Wulfnoth, although other explanations are that it was so-called to distinguish it from St Mary Woolchurch, a nearby church destroyed in the Great Fire in whose churchyard there was a beam for weighing wool, Woolnoth derived from Wool-nought. The church was rebuilt by Nicholas Hawskmoor in the early C18th. For many years the rector was Revd John Newton, friend of the poet William Cowper, and he was buried in a vault in the church in 1807. The crypt was later used as the booking hall for the new Bank station built alongside the church in 1897. In front of the church entrance is a small semi-circular paved forecourt with some planting and seating, bordered by fine railings.

Possibly so-called for being built on arches or for having arches on top of its steeple, St Mary-le-Bow was recorded here by the C11th. Destroyed in the Great Fire of 1666, the church was rebuilt by Wren between 1671-80, his designs based on the Temple of Peace in Rome. When the foundations were excavated they were thought to be those of a Roman temple but later deemed to be the crypt of the Norman church. After WWII the churchyard area was extended west and south and laid out as a garden around the statue of Captain John Smith, a member of the Cordwainers' Company, who founded the colony of Virginia. The churchyard was re-landscaped in 2008/09 and is now largely paved, with a mature plane tree and the statue of Smith remaining.

St Michael's Church is of early origin, known to have been given to the Abbey and Convent of Evesham in 1055, and in 1503 it was conveyed to the Drapers' Company. The south side of the church had a cloister, and a pulpit cross was erected in the churchyard in 1528. The church was originally open to the north to Cornhill with a small churchyard but this was sold and built over between 1547-53. The church was destroyed in the Great Fire and rebuilt by 1672, the tower initially repaired then rebuilt by 1721. Some of Wren's work was lost when St Michael's was remodelled in 1857-60 by Sir George Gilbert Scott. Many famous people were buried here, including Robert Fabian, John Stow's father and grandfather, and Sir William Cowper. The former churchyard was laid out as a garden with lawns, flower beds, shrubs and trees, and a row of benches against the church wall.

It is not known why the church was called St Nicholas Cole Abbey. The pre-Fire church was ancient, the main body predating its C14th tower and south aisle. By the C16th the church was below the surrounding ground level, which had been raised in the interim. Destroyed in the Great Fire of 1666, a new and larger church by Wren was built in 1677. The church was bombed in WWII and rebuilt in 1961/2 but later closed to worship. Gate piers and railings remain either side of the flight of steps leading up to the church door from Queen Victoria Street, with gravelled areas either side.

St Olave is the smallest intact medieval church in the City, first recorded in the late C12th. A stone church replaced the earlier building in the C13th, and the current building is primarily C15th although it was rebuilt in 1951-4. It has strong associations with the nearby Navy Office and Corporation of Trinity House. Samuel Pepys, who was buried here, called it 'Our own Church'. There was a churchyard by 1345 and well-preserved burial registers from 1563 onwards include a record of Mother Goose buried on 14 September 1586. Charles Dickens referred to it as 'My best beloved churchyard, the churchyard of St Ghastly Grim' in The Uncommercial Traveller. The entrance is via a gateway of 1658 that has a pediment with carved skull and crossbones. Until the early C20th the churchyard had many tombs set in grass but prior to 1920 it was laid out as a garden with a path from the gateway to the south door of the church, with some gravestones set against the walls.